Tigers, lions, a hippopotamus and other animals escaped from the zoo in Georgia`s capital after heavy flooding destroyed their enclosures, prompting authorities to warn residents in Tbilisi to stay inside Sunday. At least 12 people have been killed in the disaster, including three zoo workers.

An escaped hippo was cornered in one of the city`s main squares and subdued with a tranquilizer gun, the zoo said. Some other animals also have been seized, but it remained unclear how many are on the loose. Bears and wolves are also among the animals who fled from their enclosures amid the flooding from heavy rains and high winds.

There were no immediate reports that any of the fatalities were due to animal attacks. The zoo said one of the dead was Guliko Chitadze, a zookeeper who lost an arm in an attack by a tiger last month; the Interfax news agency said her husband also died in the flooding.

As of mid-afternoon Sunday, it was unclear how many animals remained on the loose or what species they are.

"Not all the animals who ran away from the zoo have been captured. Therefore, I want to ask the populace to refrain from moving about the city without" an urgent need to, mayor David Narmania said.

A full accounting of what animals were missing wasn`t immediately possible because a large part of the zoo remained underwater, zoo spokeswoman Khaati Batsilaishvili told The Associated Press.

Heavy rains and wind hit Tbilisi during the night, turning a normally small stream that runs through the hilly city into a surging river. The flooding also damaged dozens of houses.

Narmania told journalists that 12 people were known to have died.

Helicopters circled the city and volunteers and rescue workers labored to help those whose residences were damaged or destroyed, despite the potential danger from the escaped animals. About 1.1 million people live in the former Soviet republic`s capital.

The head of the Georgian Orthodox Church, Patriarch Ilia II, was quoted by the Interfax news agency as telling a Sunday Mass that Georgia`s former Communist rulers could be seen as involved in the disaster.

"When Communists came to us in this country, they ordered that all crosses and bells of the churches be melted down and the money used to build the zoo," he said. "The sin will not go without punishment. I am very sorry that Georgians fell so that a zoo was built at the expense of destroyed churches."

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